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Precision requirements are determined for space-based column-averaged CO2 dry air mole fraction (XCO2) data. These requirements result from an assessment of spatial and temporal gradients in XCO2, the relationship between XCO2 precision and surface CO2 flux uncertainties inferred from inversions of the XCO2 data, and the effects of XCO2 biases on the fidelity of CO2 flux inversions. Observational system simulation experiments and synthesis inversion modeling demonstrate that the Orbiting Carbon Observatory mission design and sampling strategy provide the means to achieve these XCO2 data precision requirements.